Date of Award

Fall 1-5-2018

Document Type


Degree Name

Master of Arts (MA)



First Advisor

Peter Moller

Second Advisor

Nicolas Mathevon

Third Advisor

Andrea Baden

Academic Program Adviser

Diana Reiss


We explored rhythmic variation in acoustic signals by geography across four species of Picoides woodpeckers. We hypothesized that there would be an effect of geographic location on the cadence of drums and rattle-type calls. No significant effect was observed. However, drum cadence was observed to significantly impact conspecific recognition in playbacks.

fig 1 drums.PNG (41 kB)
Drum cadence in Picoides

fig 2 calls.PNG (38 kB)
Call cadence in Picoides

fig 3 longitude.PNG (40 kB)
Drum cadence as a function of longitude

fig 4 duration.PNG (32 kB)
Drum duration as a function of longitude

fig 5.PNG (29 kB)
Downy drum cadence as a function of longitude

fig 6.PNG (30 kB)
Black-Backed drum cadence as a function of longitude

fig 7.PNG (31 kB)
Three-Toed drum cadence as a function of longitude

fig 8.PNG (29 kB)
Hairy drum cadence as a function of longitude

fig 9.PNG (68 kB)
Reaction time to drum to first signal

fig 10.PNG (61 kB)
Reaction time to call to first signal

fig 11.PNG (14 kB)
Interdrum interval in Downy Woodpeckers

fig 12.PNG (19 kB)
Interdrum interval in Black-Backed Woodpeckers

fig 13.PNG (12 kB)
Interdrum interval in Three-Toed Woodpeckers

fig 14.PNG (16 kB)
Interdrum interval in Hairy Woodpeckers



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